Many of the cells interior spaces, including the cytoplasm and nucleus as well as membrane-bounded organelles (e.g., mitochondria, endoplasmic reticulum, Golgi apparatus) and vesicles (e.g., endosomes, phagosomes, pinosomes, etc.), are densely packed with proteins, nucleic acids and poly-saccharide (Ellis and Minton, 2003; Minton, 2003). The human erythrocyte, for example, has a hemoglobin concentration of ~380 mg/mL. Therefore, based on hemoglobin's approximate partial specific volume (~0.74
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